Wingless insects first appeared in the Devonian period approximately 417–354 million years ago following the development of the vascular plants. They do not have a metamorphosis; instead they have an ametabolous development. They are known as Apterygota. Later the winged insects (Palaeoptera) evolved. Along with the development of the wings came a new more complex type of metamorphosis, the hemimetabolous development. Fossils of the first winged insects date back to the late Carboniferous period about 320 million years ago. They were the precursors to the modern day Odonata. The fossil record suggests that modern Odonata arose by the Triassic (237 Ma), with Anisoptera having originated at least in the Jurassic (168 Ma). All Odonata share some similar characteristics in vision, life cycle, habitat, morphology, flight, hunting and mating.